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Potential connection between helix splaying and weak lateral pressure. (a) Zinc transporter (3H90) (57), with two-helix bundles separated by a large gulf and cavities within the bundles. (b) Lipid flippase (3B60) (99), with a smaller opening between the helix bundles that may be required for protein function. In panels a and b, charged residues are displayed to delimit the hydrophobic thickness.
