| prop time spent active |
Proportion of scan samples out of all scan samples not spent resting or self-grooming (at one minute interval) |
| prop time spent outside group |
Proportion of scan samples in which animal was not in the center or periphery of the group; scans were taken twice per focal protocol: after 30 minutes and after 60 minutes |
| rate self-grooming |
Hourly rate of self-grooming bouts (bouts were considered distinct if separated by > 2 seconds) |
| rate self-scratching |
Hourly rate of self-scratching bouts (bouts were considered distinct if separated by > 2 seconds) |
| rate yawning |
Hourly rate of yawns |
| rate status display |
Hourly rate of loud call vocalization, a signal indicating dominance status [39]
|
| prop time spent grooming |
Proportion of scan samples out of all scan samples spent grooming other individuals (at one minute interval) |
| diversity grooming partners |
Diversity index of adult females grooming was received from and given to; assessed from scan samples at one minute intervals |
| diversity grooming given |
Diversity index of adult females grooming was given to; assessed from scan samples at one minute interval |
| number of female neighbors |
Absolute number of adult female neighbors within five body lengths; assessed at scans every five minutes |
| diversity female neighbors (close) |
Diversity index of adult females in close proximity (within one body length or in body contact); assessed during scans every five minutes |
| diversity female neighbors (far) |
Diversity index of adult females in proximity (within five body lengths, but further than one body length); assessed during scans every five minutes |
| diversity male neighbors (close) |
Diversity index of adult males in close proximity (within one body length or in body contact); assessed during scans every five minutes |
| diversity male neighbors (far) |
Diversity index of adult males in proximity (within five body lengths, but further than one body length); assessed during scans every five minutes |
| rate approaching males |
Hourly rate of approaching adult males within a range of five body lengths |
| rate approaching females |
Hourly rate of approaching adult females within a range of five body lengths |
| rate affiliation towards males |
Hourly rate of affiliative behavior (lip smack, mount, genital touch, friendly touch, play) directed at adult males |
| rate affiliation towards non-males |
Hourly rate of affiliative behavior (lip smack, mount, genital touch, friendly touch, play) directed at individuals other than adult males |
| rate threats towards males |
Hourly rate of threats directed at adult males |
| rate threats towards non-males |
Hourly rate of threats directed at individuals other than adult males |
| rate aggression towards males |
Hourly rate of overt aggression (bite, chase, hit) given to adult males |
| rate aggression towards non-males |
Hourly rate of overt aggression (bite, chase, hit) given to individuals other than adult males |
