Genes indicated by closed boxes on the outside of the circle are located on the positive strand, whereas those on the inside (orf1, orf2, and trnW) of the circle are located on the negative strand. The genome coordinates and GC contents are shown in the inner circle. The genes are colored based on the functional group they belong. The color scheme is shown in the middle of the circle. Four intron proteins, namely, ip1, ip2, ip3, and ip4, are drawn within the corresponding genes. Four putative clusters of tRNA genes are indicated with brackets and Roman numerals. For clarity, tRNA genes, which are present as isoforms, are indicated by attaching the anticodon to the gene name.