(a–i) Each individual was evaluated in terms of the modification types. First, the defect was classified as symmetric (S) or asymmetric (A), based on whether the modification was on a single wing (A) or on both wings (S). Next, each defect was identified as linked (L) or non-linked (N), depending on if a mutant phenotype was found on the forewing and hindwing (L) or just the hindwing (N). Finally, linked defects were classified as identical (I) or different (D), depending on whether modifications of the forewing and hindwing were identical (or similar) (I) or different (D). All individuals shown are from the F1 generation, except those in panels b and g, which are from the F2 generation. Hindwing modifications are indicated by yellow arrows. (j) The percentages of linked or non-linked individuals among all of the modified individuals. (k) The percentages of linked or non-linked individuals among all of the modified individuals, except those with LS defects. (l) The percentages of linked or non-linked individuals among LS mutants. (m) The percentages of each linkage type among all individuals with linked defects.