The increase in fish biomass (occurring inside the grey area) as a result of bottom trawling, in relation to the trawling intensity (f) and the scaled fishing mortality on benthos (m) (a,c) or the relative impact of bottom trawling on resistant compared to susceptible benthos (σ) (b,d) in a bottom-up controlled system (a,b) and a system with weak top-down control (α = 0.09, c,d). gBs = 0.3, gBr = 0.6, m = 2 for (b,d), all other parameters have default values.