Table 1.

Howls Produced in Relation to Rank

Rank	Mean number of Howls	Range
A	21.3	1–41
B	19.9	5–37.6
C	11.1	1.6–18.6
D	18	4.6–24
a	13.9	3–25.3
b	10.8	6.6–18.3

Mean number of howls (+range) produced to the different rank categories. Capital letters represent males, and lower case letters represent females.