The visually guided delayed saccade task. In the task, the monkey fixates a central dot while an oriented bar is displayed in the receptive field (RF; dashed circle) of a single V4 neuron. After a delay, the monkey is cued (by fixation spot offset) to make a saccade in 1 of 3 directions. On 2/3 of the trials, a target dot appears in 1 of 2 locations, conditions up (left) and opposite (middle), and the monkey is rewarded for making a saccade to that dot. If no target appears, the monkey is rewarded for executing a saccade to the RF stimulus (right).