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. 2013 Sep 25;8(9):e73810. doi: 10.1371/journal.pone.0073810

Table 2. Combining the contact area with the force per area measurements to calculate maximum attainable forces under different conditions.

R. pardalis S. guttatus
Friction (dry)
pads belly thigh pads belly thigh
Area at 90° (mm2) 65.1 72.4 108.1 20.1 38.8 119.1
Force per area (mN mm2) 1.5 1.0 0.5 3.0 7.3 6.9
Force (Area×Force per area) 98.3 74.5 53.0 60.9 284.1 820.3
Total force (mN) 225.8 1165
Force required (at 90°) 45.1 24.5
Friction (wet)
pads belly thigh pads belly thigh
Area at 90° (mm2) 65.1 72.4 108.1 20.1 38.8 119.1
Force per area (mN mm2) 0.2 0.3 0.3 0.1 0.1 0.3
Force (Area×Force per area) 11.1 23.9 28.1 2.2 3.9 35.7
Total force (mN) 63.1 41.8
Force required (at 90°) 45.1 24.5
Adhesion (dry)
pads belly thigh pads belly thigh
Area at 180° (mm2) 77.2 0 0 22.4 136.2 155.1
Force per area (mN mm2) 1.3 1.0 0.5 2.4 1.5 0.7
Force (Area×Force per area) 103.5 0 0 54.5 208.3 110.1
Total force (mN) 103.5 373.0
Force required (at 180°) 45.1 24.5
Adhesion (wet)
pads belly thigh pads belly thigh
Area at 180° (mm2) 77.2 0 0 22.4 136.2 155.1
Force per area (mN mm2) 0.2 0.7 0.3 0.1 0.3 0.3
Force (Area×Force per area) 17.8 0 0 2.9 39.5 46.5
Total force (mN) 17.8 88.9
Force required (at 180°) 45.1 24.5

Values are medians, as mean values were sometimes distorted by the occasional rogue value. All calculations were performed before reduction to a single decimal place.