Table 1. Environmental predictors used to derive species distribution models.

Variable theme	Variable name	Variable definition
Topography	*AspNS	Aspect  =  sin (rad (aspect))
	†AspEW	Aspect  =  cos (rad (aspect))
Climate	‡Isoth	Isothermality %
	TAR	Temperature Annual Range
	MTDQ	Mean Temperature of Driest Quarter
	MTCQ	Mean Temperature of Coldest Quarter
	RainSeasCV	Precipitation Seasonality (Coefficient of Variation)
	RainCQ	Precipitation Coldest Quarter (mm)
Land-cover	BLW	Broadleaf woodland
	ConW	Coniferous woodland
	AR	Arable
	GrassImp	Improved grassland
	GrassSN	Semi-natural grassland
	MHB	Mountain, heath, bog
	SW	Saltwater
	FW	Freshwater
	Coast	Coastal
	UrbGar	Built-up areas and gardens
Pesticides	Pest	Average number of risk hectares

^*AspNS  =  sine (radiant [aspect angle in degree]); †AspEW  =  cosine (radiant [aspect angle in degree]); ‡Isothermality %  =  Mean Diurnal Range (MDR)/Temperature Annual Range (TAR); where MDR  =  Mean of monthly (max temp – min temp)); TAR  =  Max Temperature of Warmest Month – Min Temperature of Coldest Month. Isothermality is a quantification of how large the day-to-night temperature oscillation is in comparison to the summer-to-winter oscillation. A value of 100 would represent a site where the diurnal temperature range is equal to the annual temperature range. A value of 50 would indicate a location where the diurnal temperature range is half of the annual temperature range.