In A, the cervical vertebrae of A. louisae CM 3018, modeled by subdivision surfaces (see text) are rigged to form a kinematic chain with joints at the centers of curvature of the condyles (displayed in red), with empirically-determined intervertebral separations that maximize the congruence between condyles and cotyles and associated zygaphophyseal pairs at each intervertebral joint. The articulated skeleton resembles the original specimen (B), but fortunately without the rigid steel armature. In C, a digital model of an ostrich Struthio camelus is shown in ONP, based on published data [15] of joint-by-joint intervertebral separations and flexion limits (in both mediolateral and dorsoventral flexion), and in D, an example of its extraordinary flexibility. Supplemental material: Movie S11, Movie S12, File S1.