Variation in paternity success in Oncorhynchus tshawytscha partitioned among the random effects of male, female (ovarian fluid, OF) and their interaction. The significance of each effect was tested using likelihood-ratio tests, where the log likelihood-ratio test statistic (LLR) is −2× the difference in log-likelihoods between hierarchically structured models (see main text). Significant p-values are indicated in italics.