Table 1. Model comparison for factors potentially influencing skeletal and eye abnormalities.

Fixed Effects	DF	AIC	Δ AIC
Region	9	1386	0
Refuge	133	1387	1
Year+Region	18	1393	7
Region+Species	26	1403	17
Size+Region+Species	27	1404	18
Year+Region+Species	35	1410	24
Site	463	1426	40
None	2	1429	42
Species	19	1435	48
Year	11	1437	51