In times when glucose is around, TCA-derived citrate can be converted into aceteyl-CoA by ACL, both in the nucleus and cytosol, thereby providing a nuclear/cytosolic pool of acetyl-CoA. This pool of acetyl-CoA, which seems to be independent of the mitochondrial acetyl-CoA pool, is used for histone acetylation by histone acetyltransferases, such as GCN5.