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Residue pairs k, l whose properties are within a specified range in terms of charge (p), volume (q), and hydrophobicity (r) are said to belong to the same 3D property bin φpqr (i.e., Ckl = p, q ≤ Vkl < q + Δv, r ≤ Hkl < r + ΔH). Property values for the residue pair in the hybrid that are significantly different from those observed in the protein family denote a clash.
