Table 1.

Environmental variables used to measure hypothesized environmental drivers of body size patterns in grizzly bears within Alberta, Canada

Hypothesized environmental driver and measurement variable	Units	Measurement location(s)§	Temporal scale(s)†
A. Regional habitat productivity
Temperature (Winter, Spring, Summer)	°C	home range	1971-2000
Precipitation (Winter, Spring, Summer)	mm	home range	1971-2000
Ecosystem	categories	telemetry	Ct-1
B. Inter-annual environments (deviations)
Temperature (Winter, Spring, Summer)	°C	home range	Bt-1, t0, t+1 &Ct-1, t0
Precipitation (Winter, Spring, Summer)	mm	home range	Bt-1, t0, t+1 &Ct-1, t0
C. Local habitat quality
Shrub habitat (quadratic)	%	telemetry	Ct-1
Canopy cover (quadratic)	%	telemetry	Ct-1
Variation in canopy cover	%	telemetry	Ct-1
Deciduous canopy cover (quadratic)	%	telemetry	Ct-1
Forest age (quadratic)	years	telemetry	Ct-1
Forest age variation	years	telemetry	Ct-1
Regenerating forest habitat (quadratic)	%	telemetry	Ct-1
Variation in regen. forest age	years	telemetry	Ct-1
Soil wetness (quadratic)	index	telemetry	Ct-1
D. Human footprint &activity
Private lands	%	telemetry	Ct-1
Protected area	%	telemetry	Ct-1
Mortality risk	index	telemetry	Ct-1
Safe harbour habitat	index	telemetry	Ct-1
Linear feature density	km/km2	telemetry	Ct-1
Distance to human feature	m	telemetry	Ct-1
Distance to active energy well	m	telemetry	Ct-1
E. Landscape change
Annual rate of habitat change	%	telemetry	Ct-1

§ Home ranges estimated by 50% multi-annual kernels; climate variables measured at kernel centroid; † Temporal scales relate to time of measurements; B relates to birth year &C to capture year. For inter-annual variation, 2-yrs prior to and up to 1-yr following birth or 1-yr prior to and the year of capture are considered.