Figure 2. Fitting a gene tree onto a species tree.

Gene trees are drawn with solid lines, and species trees are drawn with tubes. (a) In the case of DL (and ILS), each node x in the gene tree is mapped to (denoted by the green arrows) the most recent common ancestor (MRCA) of the species that contain gene copies descended from node x. (b) In the cases of HGT and hybridization, a smallest set of branch moves (denoted by the purple arrows) that makes the species tree identical to the gene tree and do not violate “a linear time order” is a parsimonious set of HGT or hybridization events that explain the difference between the species tree and gene tree.