The model.
(A) Migration of DNA through the agarose gel as shown by
[48; 57]. In this model, DNA undergoes cyclic
motion of alternating elongated and compact conformations, the duration of which
depends upon the size of the DNA. The elongated confirmation, in the form of
U-shaped structure, is assumed when the DNA molecule is transiently
‘hooked around’ the gel fiber. The small arrows show the
direction of DNA sliding off the hook.
(B) Breakage of the transiently trapped DNA at nicks. Nicked DNA
stays in the elongated confirmation (I) until a nick reaches the vertex at the
hook (II). Subsequent local unraveling of the DNA structure from the nick leads
to the formation of SS ends that pull apart, anchoring the DNA at the hook
(III). When the electric field-caused unraveling of the DNA duplex reaches
another nick in the opposite DNA strand, a double-strand break is formed (IV),
and the broken DNA can now slide off the hook (V).