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. 2013 Dec 18;8(12):e82757. doi: 10.1371/journal.pone.0082757

Table 2. Descriptive statistics for data used to build models to predict grizzly bear density in interior North America.

Variable Occupied areas Unoccupied areas
Inline graphic SD Min Max n Inline graphic SD Min Max n
Population size 92 96 15 765 76 0 0 0 0 14
Study area size (km2) 5110 4297 789 22875 76 6358 1629 3913 8959 14
Barren (% of study area) 6.7 8.6 0.0 34.6 76 1.7 2.1 0.1 6.7 14
Density (barren area removed) 23.0 15.1 2.5 64.6 76 0.0 0.0 0.0 0.0 14
CL relative (% of density) 1.0 0.5 0.1 1.9 76 0.8 0.3 0.5 1.0 14
Human-caused mortality (%) 3.7 3.9 0.0 20.1 76 01 0.0 0 0 14
Annual precipitation (cm) 84 40 16 199 76 52 7 43 68 14
NDVI 116 15 77 137 76 129 5 115 136 14
AET 296 104 97 443 76 342 60 228 440 14
Average annual temperature −4.0 5.0 −17 3.7 76 0.9 3.1 −5.0 4.7 14
Ruggedness 3.9 1.4 1.0 6.0 76 2.2 1.3 1.0 4.2 14
Trees (>25% per pixel) 48.1 34.7 0.0 99.5 76 69.7 30.0 8.4 99.7 14
Herb-shrub (>50% per pixel) 57.6 28.1 3.1 99.1 76 50.6 28.4 6.8 94.0 14
Salmon (% of diet) 1.0 3.0 0.0 14.0 76 1.5 3.6 0.0 10.2 14
Kokanee (% of diet) 0.7 3.5 0.0 26.0 76 0.6 1.5 0.0 5.1 14
Meat (% of diet) 25.1 15.5 0.0 58.2 76 29.3 13.9 12.5 48.1 14
Human density (humans/km2) 0.9 1.7 0.0 8.4 76 4.3 5.6 0.0 21.4 14
Livestock density (animals/km2) 1.7 5.5 0.0 39.4 76 11.5 16.9 0.0 53.2 14

Appendix S1 for detailed description of GIS derived variables. See Table S1 in

1 The kill rate in unoccupied areas was zero, but we used the mean rate of 3.7 for occupied areas during analysis so that these areas did not bias the distribution for this variable.