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(a) Function of the MnIVFeIII cofactor in catalysis by a class Ic ribonucleotide reductase: substrate reduction in α (R1) following radical initiation by MnIVFeIII cofactor in β (R2) through a radical-translocation process; (b) structure of the MnIVFeIII cofactor site in class Ic ribonucleotide reductase (x = 1 or 2, M1 = metal site 1, and M2 = metal site 2); and (c) structure of the FeIIIFeIV intermediate X in class Ia ribonucleotide reductase (x = 1 or 2).
