Results of Experiment 2, in which bees flew in a tunnel with transversely polarized illumination, as depicted by the lines on the ceiling in (a). This illumination mimics the pattern of polarized illumination that a bee would experience if it were to fly directly away from the sun, as shown in (b), or directly towards it. In (c), the thin lines (blue in the online version) depict the dance orientation histogram. The thick line (red in the online version) depicts the mean direction of the waggle axis, taking into account the up–down symmetry in the dance directions, as described in §2. This mean direction is oriented 91.7° counterclockwise relative to the horizontal rightward direction. (Online version in colour.)