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(Color online) Free-energy differences as a function of force between location 35.5 and location 40 (solid line), location 35.5 and location 45 (dashed line), location 30.5 and location 35 (dotted line), and location 50.5 and location 70 (dot-dashed line), calculated using the best-fit parameters corresponding to nucleosomes containing H4. The largest of these free-energy differences at a given force is rate limiting at that force.
