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Model of dTDP-4-keto-6-deoxyglucose bound in the FdtD cupin domain. The imidazole rings of His 221 and His 223 are located in positions ideally suited for proton transfer as indicated by the dashed lines. The carboxamide group of Gln 228 may function in stabilization of the enolate intermediate that is thought to occur during catalysis.
