Table 3. Model and parameter estimates for models explaining the variance in the relative timing deviations exhibited by Hudsonian godwits during their annual cycle (2009–2012).

Model	Parameters	Random Effects			Fixed Effects
		K	σ2	St. Dev.	Variable	β	SE	t
Departure from Beluga River	Prior Breeding Success + Prior # Nests	Ind.	2.57	1.60	Intercept	−5.23	3.05	−1.72
		Year	0.00	0.00	PBS	2.98	1.95	1.52
		Res.	32.91	5.74	P#N	3.34	1.98	1.68
Departure from Saskatchewan	Beluga River Departure	Ind.	4.92	2.22	Intercept	0.39	1.10	0.35
		Year	0.00	0.00	BRD	0.76	0.17	4.40
		Res.	52.07	7.22
Departure from Amazon	BRD + Sask. Departure	Ind.	44.60	6.68	Intercept	−0.16	1.71	−0.10
		Year	0.00	0.00	BRD	0.11	0.24	0.45
		Res.	49.74	7.05	SAD	1.02	0.16	6.21
Departure from Buenos Aires	SAD + Amazon Departure	Ind.	36.64	6.05	Intercept	1.13	1.77	0.64
		Year	0.00	0.00	SAD	0.03	0.23	0.15
		Res.	68.20	8.26	AMD	0.73	0.17	4.34
Arrival in Chiloe	AMD + Buenos Aires Departure	Ind.	1.31	1.15	Intercept	−0.39	0.68	−0.57
		Year	0.00	0.00	AMD	−0.00	0.08	−0.01
		Res.	18.74	4.33	BAD	0.97	0.07	14.5
Departure from Chiloe	BAD + Arrival in Chiloe	Ind.	3.54	1.88	Intercept	−0.25	0.53	−0.46
		Year	0.00	0.00	BAD	−0.03	0.10	−0.33
		Res.	4.16	2.04	ACH	0.05	0.09	0.55
Arrival in Beluga River	Chiloé Departure + N.A. Arr. + #Stops + Avg. Stop. Duration	Ind.	1.24	1.11	Intercept	−8.05	2.10	−3.84
		Year	0.88	0.94	CHD	0.25	0.24	1.06
		Res.	2.80	1.67	NAA	0.39	0.23	1.69
					Stops	2.19	0.51	4.33
					ASD	0.32	0.11	2.85

Bold-font variance and t-statistic values were determined to be significant at P<0.05 (n = 26).