| Palatines very short and posterior (posterior to M2) |
| Large lacrimal facial process |
| No contact between frontal and maxillary (rostrum) |
| Infraorbital foramen anterior (above P3) and small |
| Jugal extended anteriorly on the orbit at the expense of the maxillary |
| Orbit posterior |
| Zygomatic arches not strongly flared laterally |
| Wide pterygoid fossa |
| Alisphenoid canal absent |
| Postglenoid foramen medial to postglenoid process |
| Hypoglossal foramen present and double |
| Hypoglossal foramen far distal to jugular foramen |
| Occiput slightly inclined anteriorly |
| Strong nuchal and sagittal crests |
| Almond pointed shape of promontorium |
| Fenestra cochleae and aquaeductus cochleae present |
| Fenestra cochleae postero-medial to the fenestra vestibuli |
| Fenestra cochleae vertical and elongated transversely |
| Fenestra cochleae faces postero- laterally |
| Epitympanic wing of the periotic medial to the promontorium absent |
| Cochlear canal (aquaeductus cochleae) opens ventro-medially in the jugular fossa |
| Well-developed caudal tympanic process of the petrosal |
| Petrosal subarcuata fossa present, large and deep |
| Inner ear: Spiral turns of the cochlea widely separated |
| Inner ear: Subplanar coiling of the cochlea |
| Inner ear: Large anterior semicircular canal |
| Upper molars: Wide stylar shelf and presence stylar cusps (incl. D cusp) |
| Upper molars: Parastyle well-developed and mesio-labially located |
| Upper molars: Paracrista strong joining labially the parastyle |
| Upper molars: conules distinct |
| Upper molars: No hypocone |
| P3-4: Metacone absent |
| P3-4: No trace of transverse loph |
| P3-4: Simplified |
| Canine large |
| Lower molars: Paraconid present, paracristid sharp, trigonid not compressed |
| Lower jaw: No coronoid foramen |
