Bootstrap values based on 1000 pseudoreplicates are indicated at the respective nodes; scale bar indicates substitutions per site. The division of the CoVs into alpha, beta, gamma and delta groups is indicated by shaded boxes. Breda virus, a torovirus from the subfamily Torovirinae, family Coronaviridae was used as an outlier. FIPV, feline infectious peritonitis virus; TGEV, transmissible gastroenteritis CoV; BtCoV, bat CoV; PEDV, porcine epidemic diarrhea virus; SARS, severe acute respiratory syndrome; MHV, mouse hepatitis virus; PHEV, porcine hemagglutinating encephalomyelitis virus; HCoV, human CoV; BCoV, bovine CoV; TCoV, turkey CoV; IBV, infectious bronchitis virus.