Table 2.

Alternative distance metrics for quantifying the amount of evolutionary change over the two 1000-generation evolutionary trajectories shown in Figure 7. Rescaled space uses the M-matrix as a metric tensor, normalizing phenotypic change by the amount of mutational variance along a trajectory Populations 1 and 2 are those whose phenotypic optimum is at point (150, 450) and (450, 150) in phenotypic space, respectively.

	Population 1	Population 2
Euclidean distance traveled in phenotypic space	132.7	204.5
Total length of trajectory in phenotypic space	150.5	221.1
Euclidean distance traveled in rescaled space	64.8	57.9
Total length of trajectory in rescaled space	78.6	71.4