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| Variable | Meaning |
| t | Time |
| Aj(t) | Concentration of adenylate 5′ j-phosphate |
| xi(t) | Mass of the i th clone's parenchymal cells |
| x(t) | Total parenchymal mass |
| y(t) | Total mass of precursor VECs |
| z(t) | Total length of mature microvessels |
| v(t) | Tumor vascularization ( = z/x) |
