Table 3. Expected frequencies of the nine genotypes under zygotic and gametic selection in the F₂ population.

Genotype	Relative fitness fl	Zygotic selection		Gametic selection
		in fitness model	in liability model	in fitness model	in liability model
S1S1S2S2	Φ(a1+a2+i)	(1−r2)/D1	(1−r)2 f1/d1	(1−r)2/D2	(1−r)2 f1/d2
S1S1S2s2	Φ(a1+d2+j12)	2r(1−r)v2/D1	2r(1−r)f2/d1	r(1−r)(vg+1)/D2	r(1−r)(f1+f3)/d2
S1S1s2s2	Φ(a1−a2−i)	r2v1/D1	r2f3/d1	r2vg/D2	r2f3/d2
S1s1S2S2	Φ(d1+a2+j21)	2r(1−r)u2/D1	2r(1−r)f4/d1	r(1−r)(ug+1)/D2	r(1−r)(f1+f2)/d2
S1s1S2s2	Φ(d1+d2+l)	2(1−2r+2r2)x4/D1	2(1−2r+2r2)f5/d1	[(1−r)2(xg+1)+r2(ug+vg)]/D2	[(1−r)2(f1+f9)+r2(f3+f7)]/d2
S1s1s2s2	Φ(d1−a2−j21)	2r(1−r)x3/D1	2r(1−r)f6/d1	r(1−r)(vg+xg)/D2	r(1−r)(f3+f7)/d2
s1s1S2S2	Φ(−a1+a2−i)	r2u1/D1	r2f7/d1	r2ug/D2	r2f7/d2
s1s1S2s2	Φ(−a1+d2−j12)	2r(1−r)x2/D1	2r(1−r)f8/d1	r(1−r)(ug+xg)/D2	r(1−r)(f7+f9)/d2
s1s1s2s2	Φ(−a1−a2+i)	(1−r)2x1/D1	(1−r)2f9/d1	(1−r)2xg/D2	(1−r)2 f9/d2

D₁=(1−r)²(x₁+1)+2r(1−r)(u₂+v₂+x₂+x₃)+r²(u₁+v₁)+2(1−2r+2r²)x₄.

D₂=2(1−r)(x_g+1)+2r(u_g+v_g).

d₁=(1−r)²(f₁+f₉)+2r(1−r)(f₂+f₄+f₆+f₈)+r² f₃+2(1−2r+2r²)f₅.

d₂=2(1−r)(f₁+f₉)+2r(f₃+f₇).