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Illustration of the tetramer formed as a sandwich-like assembly of two hydrogen-bonded dimers of macrocyclic β-sheet peptide 2a in aqueous solution. The green arrow shows key NOEs between the layered β-sheets. Four sets of these interactions can occur in the tetramer. (For clarity, only one set is shown.) Macrocyclic β-sheet peptide 1 forms a similar sandwich-like tetramer in solution.
