Table 3. Gene expression models for the genes containing the sporulation QTN.

Gene	Gene Expression Model*	Multiple R-squared#	F-test p-value
RME1	ERME1 = −116.1+256 RME1ncV	0.8	1.9e-12
IME1	EIME1 = 394.6−506 RSF1cV−223 IME1ncV	0.6	1.8e-05
RSF1	ERSF1 = −4	0.3	0.2

*E_<gene> represents the residual expression of the particular gene after the effect of experimental variation is removed. The first term in the model (intercept) is the mean residual expression of the gene in the vineyard strain with all four oak QTN (Vineyard OOOO). Each subsequent term in the model represents the gene expression effect of replacing the oak allele of the particular QTN with the vineyard allele in the Vineyard OOOO strain (−/+ indicates direction of effect). Only significant terms in the model are shown Pr(>|t|)<0.1.

^#R-squared value obtained from applying the full model containing all possible main and interaction effects between the four sporulation QTN.