Table 1. List of symbols.

Symbol	Definition
k	Substitution rate.
Ni	No. class i individuals.
$\overline{N}$	Average no. gene copies in the population.
Pr(n)	Stationary probability that the population is in state n.
Pr(n′|n)	Forward transition probability from state n to state n′.
gi	Ploidy of a class i individual.
tij	Probability that a gene randomly sampled in a class-i individual was transmitted by a class-j individual.
μij	Mutation rate at a gene transmitted by a class-j individual to a class-i individual.
$\overline{\mu }$	Average mutation rate in the stationary demographic regime.
μe	Effective mutation rate.
μe,0	Effective mutation rate in newborns.
T	Average generation time (or mean age of the parents of a newborn).
T(n)	Average generation time in population state n.
wij(n′, n)	Expected no. class-i individuals in a population in state n′ descending from a single class-j individual in population state n.
fij(n′, n)	Probability that a gene sampled in a class-i individual when the population is in state n′ is a copy of a gene of a class-j individual given the population was in state n in the parental generation.
aij(n′, n)	Probability that a gene sampled in a class-i individual when the population is in state n′ is a copy of a gene of a class-j individual and the population was in state n in the parental generation.
πi(n)	Fixation probability of a single mutant residing in a class-i individual.
$\overline{\pi }$	Average fixation probability of a single mutant.