Associated Marker, Position refers to the marker and calculated position (cM) closest to the QTL peak. CI refers to the 2-LOD confidence interval in cM. Variance refers to the percentage of variance explained by the QTL. Additive Effect indicates the change in trait value associated with Van-0 genotypes at the indicated QTLs. Significant LOD scores were determined by calculating P values associated with the maximal LOD scores obtained by 10,000 permutations of the mapping data. Variance components were estimated by restricted maximum likelihood. Asterisks indicate QTLs that did not have a statistically significant effect on trait values by GLM.