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miRNA biogenesis in human cells. a The canonical pathway covers 1 pri-miRNA transcription, 2 Drosha cleavage, 3 pre-miRNA export to cytoplasm and 4 Dicer cleavage into 5 the miRNA/miRNA* duplex. b The alternative Drosha-independent biogenesis pathway (mirtron pathway) is indicated by the orange dashed line. See text for more details
