Cumulative SOM-derived respiration of topsoil, cryoturbated, and mineral subsoil horizons from a tundra ecosystem in the central Siberian Arctic, after incubation with glucose (Glc), amino acids (AA), cellulose (Cell) or protein (Prot). Bars represent means ± standard errors of four replicates, expressed as percent of unamended controls which are depicted by the dashed line. Significant differences between amended samples and controls are indicated (*, p < 0.05; **, p < 0.01; ***, p < 0.001).