Substrate use efficiency of topsoil, cryoturbated, and mineral subsoil horizons from a tundra ecosystem in the central Siberian Arctic, after incubation with 13C-labeled glucose (Glc), amino acids (AA), cellulose (Cell) or protein (Prot). Substrate use efficiency was estimated as the proportion of substrate-derived C in PLFAs over substrate-derived C in both PLFAs and respiration. Bars represent means ± standard errors of four replicates.