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. 2014 Jan 23;8(7):1428–1439. doi: 10.1038/ismej.2013.248

Table 3. Grouping of amino acids based on the G+C content of the first two positions in a codon.

Classification by G+C content of the first two positions in a codon
 High G+C P, A, G, R
 Low G+C F, I, M, Y, N, K, L
 Neutral V, S, T, H, Q, D, E, C, W