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. 2014 Jul;114(1):61–83. doi: 10.1093/aob/mcu081

For all sequences generated by the first author, a DNA accession number is given

Taxon Locality Voucher collector with collection no. (herbarium) DNA acc. no. GenBank accession no.
    rpl32-trnL nrITS ETS
Clinopodium dalmaticum (Benth.) Bräuchler & Heubl M. Kintgen s.n. JQ669340
Clinopodium taxifolium (Kunth) Govaerts B. Drew 228 JQ669288
Clinopodium vulgare L. Riina 1579 JQ669290
Collinsonia canadensis L. JBW 958 DQ667248
Raiche s.n. UCBG 1984.0696 JQ669291 JQ669157
Dorystaechas hastata Boiss. & Heldr. ex Benth. Anatolia Albach D6–4 (OLD) 213 KJ747319 KJ584248 KJ584275
Cult. RBGE1972–0177D JQ669302 DQ667252
HQ418845
Glechoma hederacea L. B. Drew 69 JQ669307
Horminum pyrenaicum L. Cult. RBGE 1997–2109a JQ669315 DQ667257
M. Will 63 (MJG 003069) 390 KJ747327 KJ584247 KJ584279
Hyptis laniflora Benth. B. Drew 41 JQ669317
Isolate K36721 JF301548 JF404259
Lepechinia bella Epling Bolivia P. Wester 145 (MJG 009474) 216 KJ747326
Lepechinia calycina (Benth.) Epling ex Munz Drew 197 JQ669324
Lepechinia conferta Epling Alonso 8376 (F) DQ667307
Lepechinia lamiifolia (Benth.) Epling B. Drew 178 JQ669325
Lepechinia lancifolia (Rusby) Epling Smith 444 (F) DQ667306
Lepechinia mexicana (S.Schauer) Epling B. Drew 164 JQ669326
Melissa axillaris (Benth.) Bakh.f. D. E. Boufford et al. 24526 JQ669334
Melissa officinalis L. M. Will 64 (MJG 003068) 388 KJ747325 KJ584249 KJ584285
JBW 2575 (cult. USA/WIS) DQ667291
B. Drew 70 JQ669335 JF301353
Mentha arvensis L. B. Drew 82 JQ669336
Mentha pulegium L. Riina 1574 JQ669338
Mentha spicata L. J. Walker 2566 JQ669339
Mentha spicata subsp. condensata (Briq.) Greuter & Burdet Riina 1575 JQ669337
Meriandra bengalensis (Konig ex Roxb.) Benth. Lavranus & Newton 15796 (MO 2633828) DQ667329
Perilla frutescens (L.) Britton JBW 1078 (cult. USA/WIS) DQ667246 JF301326
Perovskia atriplicifolia Benth. M. Will 65 (MJG 003070) 472 KJ584242
  JBW 2524 (cult. USA/WIS) JQ669352 DQ667223 JF301328
Perovskia scrophulariifolia Bunge Kinziraeva 6751 (MO 5201778) DQ667330
Rosmarinus officinalis L. M. Will 66 (MJG 003071) 389 KJ747310 KJ584197 KJ584296
  JBW 2558 (cult. USA/WIS) JQ669364 DQ667241
Salvia aegyptiaca L. M. Kuschewitz s.n. (cult. BG HH) 204 KJ747314 KJ584245 KJ584254
  McLeish 3728 (E) DQ667285
S. aethiopis L. Armenia J. Hellwig s.n. 26/6/02 (MJG 009919) 281 KJ584163
  Armenia J. Hellwig s.n. 26/6/02 (MJG 009919) DQ667272
S. africana-caerulea L. S Africa P. Wester & R. Claßen-Bockhoff 319 (MJG 041401) 230 KJ747271 KJ584204 KJ584255
  S Africa P. Wester & R. Claßen-Bockhoff 317 (MJG 041402) 229 KJ584203
S. africana-lutea L. S Africa P. Wester 342 (MJG 041393) 81 KJ747273 KJ584205 KJ584256
  S Africa P. Wester 708 (MJG) 234 KJ747259
  S Africa P. Wester 708 (MJG) 430 KJ747272
S. albicaulis Benth. S Africa P. Wester, R. Claßen-Bockhoff & E. v. Jaarsveld 340 (MJG 041403) 1 KJ747274 KJ584206 KJ584257
S. albicaulis × granitica S Africa P. Wester, R. Claßen-Bockhoff & E. v. Jaarsveld 341 (MJG 041404) 2 KJ747275 KJ584207 KJ584258
  S Africa P. Wester, R. Claßen-Bockhoff & E. v. Jaarsveld 341 (MJG 041404) 42 KJ584215 KJ584277
S. apiana Jeps. California P. Wester 411 (MJG 041452) 392 KJ747321
  JBW 2509 USA (WIS) DQ667214
S. areysiana Deflers Yemen Thulin, Eriksson, Gifri & Långström 8472 (UPS) 282 KJ747315 KJ584259
S. argentea L. Italy R. Claßen-Bockhoff s.n. Mai 2002 (MJG) 57 KJ747299 KJ584164
S. aristata Aucher Iran K.H. Rechinger s.n. 1974 (M) 289 KJ584244
  Iran Y. Ajani 1569 (ACECR) 417 KJ747264
  Wedelbo & Assadi s.n. (E) JQ669365 DQ667280 JF301336
S. aucheri var. canescens Boiss. & Heldr. Anatolia F. Celep 1245 (PSL METU) 239 KJ584193
  Archibald 7670 (E) DQ667286
S. aurita L.f. P. Wester & R. Claßen-Bockhoff 324 (MJG 041405) 11/424 KJ747276 KJ584218 KJ584261
  M. Will 26 (MJG 041563) 423 KJ584219 KJ584260
S. aurita var. galpinii (Skan) Hedge P. Wester 472 (MJG) 425 KJ747269
S. austriaca Jacq. R. Claßen-Bockhoff s.n. 2004 (cult. BG Mz) DQ667323
  Austria R. Claßen-Bockhoff s.n. 12.06.2003 (MJG) 280 KJ747261
S. bariensis Thulin Somalia M. Thulin, A. Dahir & A. Osman 9429 (UPS) 283 KJ747316 KJ584262
S. broussonetii Benth. M. Will 33 (MJG 041537) 29 KJ747293 KJ584226 KJ584263
  Tenerife R. Claßen-Bockhoff 2/10 (MJG 009887) 463 KJ584225 KJ584264
S. cabulica Benth. Afghanistan H. Freitag 4683 (MSB 137713) 322 KJ584189
  Ghafoor & Goodman 5148 (E) DQ667287
S. canariensis L. M. Will 46 (MJG 041565) 5 KJ747295 KJ584227 KJ584266
  Tenerife R. Claßen-Bockhoff 3/10 (MJG) 464 KJ584265
  Tenerife R. Claßen-Bockhoff 1/03 (MJG) 223 KJ747294
  Cult. RBGE 1986–0478 DQ667256
S. candelabrum Boiss. M. Will 42 (MJG 041557) 62 KJ747255 KJ584190
S. candidissima subsp. occidentalis Hedge Anatolia F. Celep 1487 (PSL METU) 201 KJ747300 KJ584165
S. candidissima Vahl. Cult. RBGE 1999–2202A DQ667261
S. chamelaeagnea Berg. P. Wester & R. Claßen-Bockhoff 314 (MJG 041407) 52 KJ747289 KJ584210 KJ584268
  S Africa P. Wester, R. Claßen-Bockhoff & E. Van Jaarsveld 313 (MJG 041406) 432 KJ584211 KJ584267
  M. Will 47 (MJG 041541) 433 KJ584212
S. chienii E. Peter M. Will 61 (MJG 003066) 53 KJ747322 KJ584250
  AnH0305–21 DQ132868
S. chionopeplica Epling JBW 2545 (cult. USA/WIS) DQ667227
S. cf. chionopeplica Epling P. Wester 485 (MJG 041435) 174 KJ747318 KJ584188 KJ584269
S. daghestanica Sosn. M. Will 34 (MJG 041551) 276 KJ747308 KJ584187
  Cult. RGB E 1988–2283A DQ667258
S. deserta Schang M. Will 96 (MJG 003100) 286 KJ747263 KJ584176
  XingJ0305–1 DQ132865
S. deserti Dcne. Egypt E. Gamal Eldin s.n. 3.5.1991 (GOET) 335 KJ747312 KJ584270
S. disermas L. P. Wester & R. Claßen-Bockhoff 326 (MJG 041413) 15 KJ747296 KJ584179 KJ584271
  M. Will 80 (MJG 003116) 454 KJ584272
S. disermas L. (syn. S. rugosa in GenBank) Goldblatt 7500 (E) DQ667290
S. dolomitica Codd P. Wester & R. Claßen-Bockhoff 321 (MJG 041411) 82 KJ747290 KJ584214 KJ584274
  F. Brusse 5610 (M) 440 KJ584213 KJ584273
  JBW 3200 (cult. USA/WIS) DQ667322
S. dominica L. Cyprus A. Seregin, D. Sokoloff & M. Remizova A-211 (MW) 267 KJ584167
  M. Kuschewitz s.n. (MJG 009323) 217 KJ747262 KJ584166
S. evansiana Hand.-Mazz. var. evansiana M. Will 55 (MJG 003060) 415 KJ747323 KJ584251
S. freyniana Bornm. Anatolia S. Bagherpour 493 (PSL METU) 98 KJ747266
S. fruticosa Miller Anatolia F. Celep 1373 (PSL METU) 100 KJ747256 KJ584195
  G. Hausner GR 31 (MJG 003078) 66 KJ584194
S. funerea Jones P. Wester 490 (MJG 041430) 393 KJ747320
S. garipensis E.Meyer ex Benth. Strohbach 149 (E) DQ667281
S. geminata Thulin Yemen M. Thulin, Beier & M. Hussein 9629 (UPS) 341 KJ584276
S. glutinosa L. Anatolia F. Celep 1196 (PSL METU) 101 KJ584253
  JBW 2568 (cult. USA/WIS) DQ667250
S. graciliramulosa Epling & Játiva Bolivia P. Wester 14 (MJG 041090) DQ667276
S. gretai Brandegee USA JBW 2511 (WIS) JQ669367 DQ667215 JF301331
S. henryi Gray USA JBW 2516 (WIS) DQ667216
S. herbanica A.Santos & M.Fernández Fuerteventura R. Claßen-Bockhoff 1/05 (MJG 009888) 40 KJ747313 KJ584246 KJ584278
S. heterochroa Fern. Yunnan, China D. E. Boufford, J. H. Chen, S. L. Kelley, R. H. Ree, H. Sun, B. Xü, J. P. Yue, L. L. Yue, D. C. Zhang & W. D. Zhu 35205 (HUH 286716) 252 KJ747324 KJ584252
S. hydrangea Benth. Anatolia A. Kahraman 1468 (PSL METU) 242 KJ747257 KJ584192
  Rechinger 47123 (E) DQ667288
S. hydrangea Benth. (syn. S. dracocepha-loides Boiss. in GenBank) Armenia Hellwig s.n. (MJG 009920) DQ667265
S. interrupta Schousb. M. Will 30 (MJG 041550) 447 KJ747265 KJ584191
S. judaica Boiss. M. Will 57 (MJG 003061) 409 KJ584241
S. lanceolata Lam. P. Wester 316 (MJG 041396) 58 KJ747278 KJ584202 KJ584281
  S Africa P. Wester 1129 (NBG) 264 KJ747277 KJ584201 KJ584280
S. lanceolata × africana-caerulea P. Wester & R. Claßen-Bockhoff 315 (MJG 041400) 236 KJ747279 KJ584216 KJ584282
S. lanigera Poir. Sinai A. El-Banhawy 11 (University of Ismailia, Egypt) 198 KJ584185
S. leucodermis Baker Madagascar R. A. Clement, P. B. Phillipson & G. Rafamantanantsoa 2137 (E 00161484) 348 KJ747280 KJ584220 KJ584284
  Madagascar P. Wester 1131 (TAN) 459 KJ584221
  Madagascar B. Bytebier 3193 (TAN) 460 KJ584222 KJ584283
S. merjamie Forsk. M. Will 83 (MJG 003113) 265 KJ747297 KJ584184 KJ584286
S. microstegia Boiss. & Bal. Armenia J. Hellwig s.n. (MJG 009884) 212 KJ747307 KJ584171
S. microstegia Boiss. & Bal. (S. verbascifolia M.Bieb.in GenBank) Armenia J. Hellwig s.n. (MJG 009884) DQ667264
S. muirii L.Bolus P. Wester & R. Claßen-Bockhoff 328 (MJG 041409) 162 KJ747283 KJ584208 KJ584287
  P. Wester & R. Claßen-Bockhoff 318 (MJG 041410) 163 KJ584209 KJ584288
S. namaensis Schinz M. Will 28 (MJG 041552) 435 KJ747284 KJ584217 KJ584290
  P. Wester & R. Claßen-Bockhoff 330 (MJG 041415) 78 KJ747281 KJ584234 KJ584291
  SW Africa W. Giess & M. Müller 14319 (M) 296 KJ584200 KJ584289
S. nilotica Juss. ex Jacq. U. Hecker g3186 (MJG 003079) 64 KJ584230
  M. Will 49 (MJG 041538) 436 KJ747258 KJ584229 KJ584292
S. officinalis L. JBW 2580 (cult. USA/WIS) DQ667225
  Voucher 2160 JQ771324
  M. Palma s.n. UCBG 7·0083 JQ669369 JF301355 JF301332
  N Africa A. El-Banhawy 16 (University of Ismailia, Egypt) 243 KJ584196
S. palaestina Benth. Anatolia F. Celep 1083 (PSL METU) 400 KJ747304 KJ584175
  Anatolia A. Kahraman 1443 (PSL METU) 125 KJ747301
S. cf. palaestina Benth. N Africa A. El-Banhawy 6 (University of Ismailia, Egypt) 200 KJ747302 KJ584172
S. patens Cav. Cult RBGE 1973–9197 JQ669370 DQ667253 JF301333
S. penstemonoides Kunth & Bouché JBW 2578 (cult. USA/WIS) DQ667221
S. phlomoides ssp. phlomoides Asso Morocco R. Vogt 10336 & Ch. Oberprieler 4784 (B 100145114) 337 KJ747309 KJ584186
S. polystachya Ort. Breedlove & Mahoney 72286 (UC) cult. UCBG 92·052 JQ669371 JF301334
S. pratensis L. C Russia A. Suchorukow s.n. VII 2003 (MW) 275 KJ747291 KJ584178
  Isolate S0628 EU169486
S. przewalskii Maxim. Cult. RBGE 1993–2067A JQ669372 DQ667254 JF301339
S. radula Benth. S Africa Germishuizen 3950 (MO 4385830) 328 KJ584180 KJ584293
S. repens Burch. ex Benth. M. Will 50 (MJG 041931) 61 KJ747282 KJ584232 KJ584295
S. cf. repens Burch. ex Benth. P. Wester & R. Claßen-Bockhoff 325 (MJG 041412) 437 KJ584231 KJ584294
S. roemeriana Scheele USA JBW 2515 (WIS) DQ667211
S. scabra L. M. Will 37 (MJG 041549) 55 KJ747285 KJ584233 KJ584297
S. schimperi Benth. Yemen D. Podlech 36057 (M 55003) 310 KJ584174 KJ584298
  M. Will 72 (MJG 003099) 466 KJ584168
S. schlechteri Briq. P. Wester & R. Claßen-Bockhoff 323a (MJG 041416) 31 KJ747286 KJ584235 KJ584299
  M. Will 51 (MJG 041539) 438 KJ584236 KJ584300
S. sclarea L. Anatolia F. Celep 1492 (PSL METU) 244 KJ747305 KJ584162 KJ584301
  JBW 2527 (cult. USA/WIS) JQ669373 DQ667222
S. sessilifolia Baker Madagascar R. A. Clement, P. B. Phillipson & G. Rafamantanantsoa 2001 (MO 4328854) 331 KJ584223 KJ584302
  Madagascar C. H. Jongkind & S. Rapanarivo 929 (MO 4870099) 332 KJ584224 KJ584303
  Madagascar Jongkind & Rapanarivo 929 (E) DQ667282
S. somalensis Vatke M. Will 77 (MJG 003119) 340 KJ747311 KJ584240 KJ584304
S. spinosa L. N Africa A. El-Banhawy 14 (University of Ismailia, Egypt) 199 KJ747303 KJ584173
S. stenophylla Burch. ex Benth. S Africa Burgoyne & Snow 4805 (MO 5649981) 330 KJ747260 KJ584237 KJ584305
  Giess & Hübsch 11607 (M) 439 KJ584238 KJ584306
S. summa A. Nelson Texas JBW 1972 USA (WIS) DQ667217
  Texas P. Wester 373 (MJG 041338) 190 KJ584307
S. sylvestris L. Anatolia A. Kahraman 1568 (PSL METU) 118 KJ747292 KJ584181
(syn. S. tesquicola Kiok. & Pobed.) C Russia A. Suchorukow s.n. VIII 1994 (MW) 273 KJ584177
  Isolate S0626 EU169485
S. taraxacifolia Hook.f. Morocco W. Lippert 25355 (M) 304 KJ747270 KJ584228 KJ584308
  JBW 2521 (cult. USA/WIS) DQ667209
S. texana (Scheele) Torr. Texas P. Wester 362 (MJG 041477) 191 KJ747267 KJ584199
  Texas P. Wester 362 (MJG 041477) DQ667321
S. thermarum Van Jaarsv. P. Wester & R. Claßen-Bockhoff 336 (MJG 041398) 23 KJ747288 KJ584239 KJ584309
  M. Will 52 (MJG 041933) 452 KJ584310
  S Africa P. Wester 312 (MJG 041397) 221 KJ747287
S. tingitana Etling Morocco D. Podlech 43384 (M 54979) 314 KJ584169
S. trichocalycina Benth. Breckle 4963 (E) DQ667283
S. trijuga Diels YunN0309–5 DQ132870
S. veneris Hedge Cyprus M. J. Y. Foley 1701 (E 00147797) 413 KJ747306 KJ584170
S. verbenaca L. Syria W. Licht SYR 307 (MJG 003082) 67 KJ584182
  Anatolia F. Celep 1408 (PSL METU) 141 KJ747298 KJ584183
S. whitehousei Alziar Texas P. Wester 352 (MJG 041389) 231 KJ747268 KJ584198 KJ584311
  Texas P. Wester 352 (MJG 041389) DQ667320
Thymus pulegioides L. Riina 1577 JQ669378
Thymus serpyllum L. J. Walker 2564 (cult. USA/WIS) JQ669379
Zhumeria majdae Rech.f. & Wendelbo Iran F. Sharififar 1651 (ACECR) 422 KJ747317 KJ584243 KJ584312
  Wendelbo 15793 (V 21730) DQ667336
  Ghazi s.n. (V 01176) DQ667335

acc. no., accession number; locality: origin from natural habitats confirmed; ACECR, Iranian Academic Center for Education Culture and Research; BG HH, Botanical Garden Hamburg (Germany); cult., cultivated; PSL METU, Plant Systematics Lab. Department of Biological Sciences, Middle East Technical University, Ankara (Turkey); RBGE, Royal Botanical Garden Edinburgh (UK); herbarium acronyms according to Index Herbariorum.