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. Author manuscript; available in PMC: 2015 Jan 21.
Published in final edited form as: Chem Soc Rev. 2014 Jan 21;43(2):676–706. doi: 10.1039/c3cs60249d

Table 7.

The nap operon gene content and organization in bacteria

Quinone oxidase nap Operon Phylogenic class Organism
H AGHBF_LD ε Sulfurovum sp. Autotrophic
AGHBFLD Aquificae Hydrogenivirga sp. 128-5-R1-1
Persephonella marina EX-H1
Sulfurihydrogenibium sp. YO3AOP1
Deinococci Oceanithermus profundus DSM 14977
ε Arcobacter (butzleri RM4018 and Nitrofigilis) Campylobacter species
(concisis, curvis, hominis, fetus)
Helicobacter mustelae 1219
Nautilla profundicola
Sulfuricurvum kujiensedum
Sulfurospirillum (barnesii and deleyianum)
Wolinella succinogenes
AGHBFLDS ε Nitratiruptor sp. SB155-2
Caminibacter mediatlanticus
AGHBFLSD ε Thiomicrospira (formerly Sulfurimonas) denitrificans DSM 125
AGHBLD ε Campylobacter (lari, coli, jejuni (8 strains), upsalensis) Helicobacter hepaticus
DAGHB β. Burkholderiales bacterium 1_1_47
γ Shewanella amazonensis SB2B*, halifaxensis HAW-EB4, loihica PV-4*, oneidensis MR-1,
pealeana ATCC 700345*, sediminis HAW-EB3*, sp. ANA-3*, MR-4*, MR-7, W3-18-1*
DAGHB_F< β Candidatus Accumulibacter phophatis
DAGHF δ Geobacter lovleyi
DGBAH:G Clostridia Desulfitobacterium hafniense strains (2 DCB-2 & 2 DCB-1)
FDAGHB α Magnetospirillum magnetotacticum AMB-1
GBAH:G Clostridia Desulfitobacterium hafniense Y51
MADGH Chrysiogenetes Desulfurispirillum indicum
Deferribacteres Calditerrivibrio nitroreducens
Deferribacter desulfuricans
Denitrovibrio acetiphilus
C ABC γ Photobacterium (damselae CIP 102761 and profundum SS9*)
AC Sphingobacteria Rhodothermus marinus
CDAG Clostridia Symbiobacterium thermophilum
DABC β Dechloromonas aromatica RCB
γ Hahella chejuensis KCTC 2396
Photobacterium sp. SKA34*
Shewanella (piezotolerans WP3*, sediminis HAW-EB3*, violacea DSS12)
Vibrio vulnificus CMCP6
DABCF Flavobacteria Maribacter Sp. HTCC2170
EABC γ Photobacterium profundum 3TCK*
EDABC α Afipia sp. 1NLS
Azospirillum sp. B510
Bradyrhizobium japonicum USDA 110
Bradyrhizobium sp. BTAi1
Bradyrhizobium sp. ORS278
Methylobacterium sp. 4-46
Paracoccus (denitrificans PD1222 and pantatrophus) Rhodopseudomonas palustris BisB18
β Achromobacter xylosoxidans A8
Azoarcus sp. BH72*
Burkholderia xenovorans LB400
Comamonas testosteroni CNB-2
Cupriavidus metallidurans CH34, taiwanensis LMG 19424 and necator
(formerly Ralstonia eutropha H1 6)
Ralstonia (eutropha JMP134 and pickettii strains 12D & 12J)
γ Marine γ proteobacterium HTCC2148
Pseudomonas (mendocina ymp, and stutzeri A1501)
Saccharophagus degradans 2-40
Shewanella amazonensis SB2B*, baltica OS155, baltica OS185*, baltica OS195*,
baltica OS223, denitrificans OS217, frigidimarina NCIMB 40, loihica PV-4*,
pealeana ATCC 700345*, ANA-3*, MR-4*, W3-18-1*, woodyi ATCC 51908*
EFDABC α Agrobacterium tumefaciens str. C58
Pseudomonas sp. G-179
Rhizobium leguminosarum bv. trifolii WSM2304
Rhodobacterales bacterium Y4I
Rhodospirillum centenum SW
Sinorhizobium (fredii NGR234, medicae WSM419, meliloti 1021)
Starkeya novella
γ Colwellia psychrerythraea 34H
Pseudomonas aeruginosa (strains LESB58, UCBPP-PA14, PAO1 & PA7)
Reinekea sp. MED297
FDABC γ Pantoea sp. At-9b, Photobacterium (angustum S14, profundum strains (3TCK* & SS9*))
Psychromonas ingrahamii 37
Serratia odorifera 4Rx13, proteamaculans 568, Vibrio (cholerae str. N16961,
harveyi ATCC BAA-1116,fischeri strains (ES114, MJ11), parahemeolyticus
RIMD 2210633, vulnificus YJ016)
Yersinia (pestis (5 strains), enterocolitica)
FDABC_ABC γ Moritella sp. PE36
KEFDABC α Rhodobacter sphaeroides strains (2.4.1 & ATCC 17025*)
HC CMADGH Aquificae Thermovibrio ammonificans
δ Desulfovibrio desulfuricans strain 27774
Desulfovibrio sp. 3 1 syn3
DAGHBC β Leptothrix cholodnii
γ Aggregatibacter aphrophilus
Citrobacter (koseri and rodentium)
Escherichia coli IAI3
Salmonella typhimurium
Shigella flexneri (2 strains)
FDABGHBC α Rhodobacter sphaeroides ATCC 17025
FDAGHBC α Dinoroseobacter shibae DFL 12
Laribacter hongkongensis
β Azoarcus sp. BH72*
Thauera sp. MZ1T
γ Aeromonas hydrophila
Edwardsiella (ictaluri, tarda)
Escherichia coli (16 strains), Grimontia hollisae CIP 101886
Haemophilus influenza
Pectobacterium (atrosepticum, carotovorum (2 strains), wasabiae)
Providencia (alcalifaciens, rettgeri, rustigiianii, stuartii)
Salmonella enterica
Serratia odorifera DSM 4582
Shigella (boydii (2 strains) and dysenteriae)
NA A γ Photobacterium sp. SKA34*
ε Helicobacter felis
AB δ Sorangium cellulosum
DAB γ Shewanella (baltica OS185*, baltica OS195*, Piezotolerans WP3*, woodyi ATCC 51908*)
MA δ Anaeromyxobacter sp. K and dehalogenans strains (2CP-1 & 2CP-C)

The nap operons categorized into four groups based on the presence or absence of a membrane quinone oxidase (napC or napH). The gene order in each organisms nap operon was provided by the NCBI gene and phenotype mapping database (http://www.ncbi.nlm.nih.gov/gene), and in a few cases the JGI database (http://img.jgi.doe.gov). The identity of each translated Nap protein was confirmed by performing NCBI database blast searches. In each operon the nap genes are listed by the order in which they are found. Organisms that encode two separate nap operons are shown with asterisks (*); fused operons with a colon (:), underscore indicates a gap, and gene encoded on the complementary strand is indicated with an arrow (<). Proteobacteria are denoted as α, β, γ δ, ε.