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The distribution of dIV intrinsic muscles in (a) the primary anisodactyl quail, (b) the zygodactyl budgerigar and (c) the putative secondary anisodactyl zebra finch. Budgerigar lost the EBDIV but conserves the ABDIV. Both muscles are absent in the zebra finch. (Online version in colour. Muscles are shown in red, skeleton in blue.)
