Table 3. Results of DISTLM procedure for fitting environmental variables to macrobenthic composition.

Marginal tests			Sequential tests
variable	psF	R2	variable	psF	R2	cumR2
month	4.6**	0.16	bottom type	38.0**	0.28	0.28
site	15.5**	0.14	month	7.0**	0.16	0.44
bottom type	38.0**	0.28	site	30.3**	0.14	0.58
POC/N	2.3	0.02	POC	2.4*	0.01	0.59
POC	8.1**	0.08	mean grain size	1.9	0.01	0.60
Chla	9.8**	0.09	fine sand	1.7	0.01	0.61
mean grain size	4.8**	0.05	Chla	1.0	0.00	0.61
fine sand	5.2**	0.05
δ13C	5.2**	0.05

Analyses based on double root transformed macrofaunal data of species abundances in samples. Significant effects: * P<0.05; ** P<0.001.