Table 2.

Partitioning of total variance in LRS (in %) into its main components for 3 samples: 1) breeders, 2) sexually mature (≥4 years old), and 3) all subjects based on Brown’s (1988) equation

Components	Males			Females
	Breeders	Sexually mature	All subjects	Breeders	Sexually mature	All subjects
Breeders	100.0	83.6	57.3	100.0	81.6	27.1
Longevity (L)	26.0a	21.7	14.9	64.9b	52.9b	17.6
Fecundity (F)	62.9b	52.6b	36.1a	7.6	6.2	2.1
Q LF	11.0	9.2	6.3	27.5a	22.4	7.5
Nonbreeders	—	16.4	42.7	—	18.4	72.9
Matures	—	16.4	4.6	—	18.4	2.9
Immatures	—	—	38.1a	—	—	70.0b

The total variance due to breeders is divided into the contribution of longevity (L), fecundity (F), and their interaction (Q) (see Methods for details). Contribution of nonbreeders (i.e., subjects that did not reproduce during their lifetime) is divided into the contribution of subjects that died before reaching maturity (i.e., “immatures”) and those that reached sexual maturity (i.e., “matures”).

^aFactors explaining ≥25% of the variance.

^bFactors explaining ≥50% of the variance.