HD-ZIPIII genes are required for integument morphogenesis. (a)–(f) cna phb phv triple mutant ovules; (g)–(i) phv-1d gain-of-function ovules. (a)–(d) whole-mount ovule clearings. (e)–(i) SEMs. Scale bar = 20 μm in (a)–(d), (i). Scale bar = 50 μm in (e), (f), (h). Scale bar = 5 μm in G. Abbreviations: nucellus (n), outer integument (oi), inner integument (ii), funiculus (f), amorphous integument (ai), integument (i).