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Schematic illustrating the proposed circuit organization. L6 CT neurons provide strong input to L5a pyramidal neurons, as well as FS cells in L4 and L5. Somatostatin-expressing inhibitory interneurons and L4 excitatory neurons receive weak L6 CT input. The net effect of this synaptic organization is that L6 CT activation evokes action potentials in L5a pyramidal neurons while suppressing L4 excitatory cells.
