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(A) GFP-fusion of Umbreamel-tails with HP1B-CD (green) localizes to centromeres (anti-Cid in magenta, bar = 5 microns, colocalization appears white). (B-D) D. melanogaster Kc cell centromeric localization (Cid, red) of Umbrea orthologs (GFP, green) from D. simulans, D. teissieri and D. yakuba worsens with increased divergence. (E) Steps to essential neofunctionalization by Umbrea following gene duplication.
