There are errors in Equation (3). In the description of Equation (3), the equilibrium frequencies PIX are incorrectly defined in terms of pi, where in fact they should be defined in terms of phi. Please see the corrected description of Equation (3) here.
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Reference
- 1. Kosakovsky Pond S, Delport W, Muse SV, Scheffler K (2010) Correcting the Bias of Empirical Frequency Parameter Estimators in Codon Models. PLoS ONE 5(7): e11230 doi:10.1371/journal.pone.0011230 [DOI] [PMC free article] [PubMed] [Google Scholar]