Mutation of HCCS Domain I residues alter heme interactions.
Recombinant GST-HCCS protein (alone) and GST-HCCS: cytochrome c cocomplexes
were purified from Δccm E. coli and prepared
for UV/vis absorption spectroscopy and SDS-PAGE. Shown are spectra
for (A) WT HCCS/cyt c, (B) W118A HCCS/cyt c, and (C) N128A/M130A HCCS/cyt
c following purification (black line), chemical reduction with sodium
dithionite (red), and extraction with pyridine (inset). (D) Soret
peak spectra were obtained from cocomplexes representing WT HCCS/cyt
c (left) and W118A HCCS/cyt c (right) following purification (black)
and treatment with 100 mM imidazole (purple). UV/vis spectra between
500–580 nm (alpha/beta region) of cocomplexes treated with
100 mM imidazole following chemical reduction with sodium dithionite
are shown for (E) WT HCCS/cyt c, (F) W118A HCCS/cyt c, (G) N128A/M130A
HCCS/cyt c, (H) N128A HCCS/cyt c, and (I) M130A HCCS/cyt c. (J) Heme
stain of the indicated purified cocomplexes following SDS-PAGE and
transfer to nitrocellulose. (K) Heme stain (top) and GST-HCCS immunoblot
(bottom) of the indicated purified cocomplexes following SDS-PAGE
and transfer to nitrocellulose. (L) UV–vis spectra of sodium
dithionite reduced purified cocomplexes from WT HCCS (black), Y120A
HCCS (red), and P121A HCCS (blue). (M) UV–vis spectra of HCCS
proteins (alone) purified from WT HCCS (black) and Y120A/P121A HCCS
(green). (N) UV–vis spectra of purified cocomplexes from WT
HCCS/cyt c (black) and Y120A/P121A HCCS/cyt c (orange). (O) Heme stain
(top) and GST-HCCS immunoblot (bottom) of the indicated purified cocomplexes
following SDS-PAGE and transfer to nitrocellulose. (P) UV–vis
spectra of HCCS proteins (alone) purified from WT HCCS (black), Y120A
HCCS (red), and P121A HCCS (blue). Arrows indicate wavelength (nm)
of peak absorption maxima. All spectra were performed with equal amounts
(50–100 μg) of total purified protein. All SDS-PAGE samples
were equally loaded (2–5 μg of total purified protein
each). For all proteins, Bradford quantitation was confirmed by Coomassie
staining, which also indicated that GST-HCCS proteins were obtained
at >90% purity.