Outward adaptation of target-directed microsaccades. A, Experimental procedure. The observer initially fixated the central dot target (1). After a random delay of 500–700 ms, the dot was displaced 20 arcmin to the right (2). During the saccade, the target was shifted by 15 arcmin to the right (3) such that the target was to the right of the gaze direction after the saccade (4). B, C, Example traces of horizontal eye position. D, Mean saccade amplitudes and SDs before (green) and after (blue) saccadic adaptation for each of the six participating subjects. E, Means across subjects. Error bars indicate SEM. *p < 0.05 (one-sample two-tailed t test).