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MA1 is the best surrogate for ANE for which we have data.
Europeans share more alleles with MA1 than with Karitiana, as we see from the fact that in a plot of f4(Test, BedouinB; MA1, Chimp) and f4(Test, BedouinB; Karitiana, Chimp), the European cline deviates in the direction of MA1, rather than Karitiana (the slope is >1 and European populations are above the line indicating equality of these two statistics).
