Virus symptom severity was assessed the day before each experiment was terminated by rating each leaf by visual estimation of the proportion of the leaf showing symptoms using a 0 to 3 scale in which: 0 = no symptoms, 1 = <1/3 of the leaf involved, 2 = 1/3–2/3 of leaf area involved, and 3 = >2/3 of the leaf involved. + N = nitrogen provided as KNO3, - N = no nitrogen provided. V0 = plants derived from non-inoculated, virus-tested plant stock; V1 = plants derived from V0 plant stock graft inoculated with the potyvirus complex (SPFMV, SPVG, SPVC, and SPV2); V2 = plants derived from plant stock infected with SPCSV. Severity ratings were transformed using log 10 and Fisher's LSD test at the 0.05 probability level was used to test for statistical significance. The data are expressed as means ± SE from non-transformed data.