| γG
|
Effect of G allele at susceptibility locus |
| γg
|
Effect of g allele at susceptibility locus |
|
ϕF
|
Effect of F allele at infectivity locus |
|
ϕf
|
Effect of f allele at infectivity locus |
|
pg
|
Frequency of the g allele for susceptibility |
|
pf
|
Frequency of the f allele for infectivity |
 |
Average individual susceptibility |
 |
Average individual infectivity |
|
rγ
|
Relatedness at susceptibility locus |
|
rϕ
|
Relatedness at infectivity locus |
|
βij
|
Pairwise transmission rate parameter between susceptible individual i and infective individual j
|
| α |
Rate of recovery parameter |
| c |
Contact rate |
|
R0
|
Basic reproduction ratio |
 |
Breeding value for R0 of individual i
|
 |
Additive standard deviation in total breeding value |
| D |
Measure of linkage disequilibrium |
|
FIS
|
Measure of deviation from Hardy–Weinberg equilibrium |
